Rather than integrating different series into one or two unified universes like previous titles of the series, A.C.E.: R presents each featured series as independent timelines, with the series' main characters proceeding with their own stories before the events of the game.
The main events of the game itself revolve around an original timeline again completely independent of all other featured series. In this timeline humanity has expanded beyond the Solar system, with planet Area becoming one of those resulting colonies. But somewhere along the line the Seasons, androids originally built to help the colonists settling on Area, rebelled against their human companions. Leading a robotic army collectively known as the Icon, the Seasons successfully wiped out the humans on planet Area, but there is one last trick up the colonists' sleeves: they made a data copy out of one of the Seasons androids and entrusted it, named Autumn-4, with genetic seeds of themselves as well as a trans-atmos
